Carbon use and calcification strategies in a diversity of fleshy and calcifying tropical algae across irradiance gradients on reefs of Little Cayman Island

Marguerite S. Koch and Conall McNicholl

1Biological Science Department, Florida Atlantic University, Boca Raton, Florida, USA

Correspondence: Marguerite S. Koch, mkoch@fau.edu

Tropical reef macroalgae in varying light environments may be differentially utilizing CO2 versus HCO3 for photosynthesis. We hypothesized that macroalgae in low-light environments utilize CO2 due to energetic costs of taking up HCO3. Further, we examined calcification strategies across irradiance gradients. Fleshy and calcified macroalgal species were collected (~200 individuals; 3 Phyla) from a shallow high-irradiance barrier reef (~1,500 μmol photons m-2 s-1; ~1 m), 2 mid-depth intermediate-irradiance reefs (~600 μmol photons m-2 s-1; ~17 m), 2 deeper wall reefs (~400 μmol photons m-2 s-1; ~25 m), and low-irradiance crevices (~15 μmol photons m-2 s-1; >15 m). The 13 δC of organic and inorganic tissue were analysed. The 15 δN, total N and C were also determined, as well as water column nutrients and irradiance. The 13 δC of organic tissue ranged from -11.8 to -25.5 with an average of -19.1 ± 3.2, indicating that all species likely utilized HCO3. Ochrophytes had the most enriched organic 13δC signatures (-16.5), while chlorophytes and rhodophytes were similarly lighter (-20.6 and -20.9, respectively). Irradiance was a strong predictor of organic 13 δC (13 δC = 3.6 x 10-3 irradiance – 21.5; R2 = 0.91). Inorganic 13 δC of calcifiers was genus-specific with the chlorophytes ranging between 0.5 to 3.5, ochrophytes 2.9 to 5.2, and rhodophytes 1.7 to 3.4, except for Jania and Amphiroa that had negative signatures between -2.8 to -5.1. These data lead us to suggest that tropical macroalgae primarily utilize HCO3 for photosynthesis, with some constraints from irradiance, but taxonomic differences likely drive physiological mechanisms for calcification.